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- Cuticle annulated, without distinctly annulated internal
layer; annuli smooth.
- Lateral field with two wings (three incisures).
- Lip region weakly offset, crown-shaped, consisting of
six leaf-like lips equal in size. Pairs of lips separated by
primary axils and secondary axils without guard processes,
U-shaped clefts of axils internally sclerotized.
- Six outer labial and four cephalic papilliform sensilla.
Cephalic probolae trilobed, with median rectangular lobe and
lateral triangular lobes. Labial probolae absent. .
- Amphidial aperture rounded, located on lateral lips.
- Stoma divided into cheilostom - barrel-shaped with strongly
sclerotized bacilliform cheilorhabdia; gymnostom - barrel-shaped,
as wide as stegostom, with strongly sclerotized plate-like
gymnorhabdia; and stegostom with funnel-shaped prostegostom and
variably shaped mesostego-, metastego- and telostegostom parts.
- Stoma not armed, no metastomatal denticle .
- Pharynx cephaloboid:; isthmus narrower than metacorpus; basal
pharyngeal bulb oval, with strongly developed valves.
- Nerve ring encircling posterior part of metacorpus. Excretory
pore opens at level of nerve ring, at around the level of
metacorpus or metacorpus-isthmus junction. Deirids present.
Female:
- Reproductive system cephaloboid; posterior part of ovary
straight; spermatheca present or reduced; postvulval uterine sac
present; vulva protruding, flat or in a depression; vagina
straight.
- Phasmids located at level of anterior part of the tail
- Female tail conoid, straight, tail terminus with pointed or
harpoon-shaped mucro;
Male:
- Reproductive system cephaloboid; spicules cephaloboid with
manubrium and corpus of approximately equal width
- Gubernaculum plate-like; cornua crurum absent.
- One or two ventrosublateral pairs of genital papillae located
anterior to cloaca; one ventrosublateral pair located at the level
of cloacal opening; two pars located at middle of tail length; and
three pairs (lateral, subventral and subdorsal) near tail
terminus; there is a midventral papilla on anterior cloacal lip.
- Rectum short (1.1-1.8 times longer than anal body diameter).
- Phasmids located at around mid-tail in males. male tail
conoid, arcuate ventrad, tail terminus with pointed mucro.
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