Males:
- Bursa asymmetrical: dorsoventral height larger in right lobe
and lateral width much larger in left lobe, and dorsal and ventral
incisions present; Ray 1 minute, at level of gubernaculum). Bursal
rays characteristic of species (see refernce source for details)..
- Spicules equal length, alate, joined at distal ends and
abruptly bent dorsally, about 14% of body length
Ref: Hasegawa and Syaffrudin, 1997
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