Female: Cyst stage present; circumfenestrate;
eggs without punctation or tubercles.
Cysts are lemon-shaped, dark brown to black color, 460-840 µm
length, with a length/width ratio of 1.2-2.1, strongly striated
pattern on the cuticle.
Prominent vulval cone with circumfenestrate vulva, and an anus
located 25-60 µm from the fenestra in a depression in the cuticle.
Egg surfaces bear conspicuous punctations.