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Refer to subfamily diagnosis (Tylenchinae).
Female:
- Body straight or slightly ventrally curved after fixation.
- Cuticle with prominent annuli with zigzag margins.
- Lateral field conspicuous, with two smooth incisures
- Cephalic region continuous with body contour, hemispherical,
with very fine annuli and a small ellipsoid-hexagonal labial plate
- Amphid apertures crescent shaped,
- Stylet delicate, conus forming 28-42% of its total length, knobs
small, rounded and slightly posteriorly sloping.
- Dorsal esophageal gland outlet at 1.5 μm from stylet base.
- Procorpus elongate, cylindrical, metacorpus ovate,with distinct
central valva. Isthmus slender and long, postcorpus elongate-saccate
and offset. Cardia rounded.
- Excretory pore at posterior half of isthmus.
- Reproductive system monodelphic-prodelphic, outstretched
ovary with oocytes in single row, short oviduct, and an offset, oval
to rounded spermatheca containing spheroid sperm. Crustaformeria
quadricolumellate.
- Vagina slightly anteriorly directed, vaginal walls slightly
thickened, vulva a simple transverse slit, slightly sunken into body
and lacking flaps.
- Post-uterine sac absent.
- Tail elongate conoid, gradually tapering to a pointed to sharp
terminus bearing several small bristles on tip.
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Males:
- General morphology similar to that of female
- Testis single and outstretched.
- Spicules tylenchoid, slender, manubrium slightly offset, shaft
remarkably long and slender, blade ventrally arcuate at distal
region.
- Gubernaculum trough-shaped.
- Cloacal lips projecting, forming a short, posteriorly directed
penial tube.
- Bursa absent.
- Tail similar to that of female.
Ref. Panahandeh et al., 2019
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