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Males:
- Spicules slightly S-shaped without central lamella
- Gubernaculum with lateral flanges without denticles.
- Precloacal supplements papilliform, cup-shaped or
button-shaped, cuticle usually strongly annulated between precloacal
supplements. Type A (Weiser and Hopper, 1967)
Ref: Ott, 1972; Chu et al., 2023; Tchesunov, 2013
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