Rev 03/09/2021

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      Oriverutus Siddiqi, 1971
(The name Mammillonema a reference to the nipple-like form of the labial and cephalic papillae.)

The genus Oriverutus Siddiqi, 1971 has around 35 valid species as of 2014. In geographical distribution  its representatives mainly inhabit tropical areas of at least three continents (Africa, America and Asia) and two islands or archipelagos (Papua New Guinea and Mauritius); it is occasionally found in temperate regions (Korea, Japan, Spain) (Pena-Santiago et al., 2014)..

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Morphology and Anatomy:

Morphologically, Oriverutus is a well-characterized genus, distinguishable by a peculiar combination of features: lobe-like lips, odontostyle relatively long and slender with short aperture, pharyngo-intestinal junction often bearing three cardiac cells, and tail conically elongated usually with more or less (but always perceptible) bent dorsad end.

  • Small to medium length, 0.55 - 2.21 mm.
  • Cephalic region offset and slightly expanded from body contour.
  • Lips conspicuously separated with protruding button-like labial and cephalic papillae.
  • Amphid apertures large, occupying about three-quarters of the lip region diameter.
  • Odontostyle long (1.2 - 2.5 x lip width) and slender, aperture small.
  • Odontophore rod-like.
  • Esophagus dorylaimoid, basal expansion 0.4 to 0.5 of total esophagus length; cardia valve-like.
  • Tail region conical. often curved ventrally, similar in both sexes..


  • Of the species known in 2014, 13 are monodelphic-opisthodelphic, the remained are didelphic-amphidelphic.


  • Males only known for one monodelphic-opistyhodelphic species, but common in the didelphic species
  • one to 11 pairs of equally spaced ventromedian supplements

Ref: Pena-Santiago et al (2014).

Body size range for the species of this genus in the database - Click:

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 (Yeates et al., 1993).

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Biology and Ecology:

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Life Cycle:

For Ecophysiological Parameters for this genus, click 
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Ecosystem Functions and Services:


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Ahmad, W. 2007. New and known species of Dorylaimoidea (Nematoda: Dorylaimida) from Singapore. Nematology 9:215-229.

Andrássy, I. 2009.  Free-living nematodes of Hungary, III.  Hungarian Natural History Museum. 608p.

Liebanas, G., and Pena-Santiago, R. 2011. Nematology 13:463-475.

Liebanas, G., Esquivel, A., and Pena-Santiago, R. 2011. Nematology 13:307-318. 

Pena-Santiago, R. Liebanas. G. and Ahmad, W. 2014. Taxonomy of the genus Oriverutus Siddiqi, 1971 (Nematoda: Dorylaimida: Nordiidae). Journal of Natural History 48:21-22, 1253-1278, DOI: 10.1080/00222933.2013.861942 .

Siddiqi MR. 1971. Oriverutus lobatus gen. n., sp. n. and Sicaguttur sartum gen. n., sp. n. (Nematoda: Dorylaimoidea) from cultivated soils in Africa. Nematologica. 16:483-491.

Yeates, G.W., T. Bongers, R. G. M. De Goede, D. W. Freckman, and S. S. Georgieva. 1993. Feeding habits in soil nematode families and genera—An outline for soil ecologists. Journal of Nematology 25:315-331.


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Copyright 1999 by Howard Ferris.
Revised: March 09, 2021.